Every mob in the open-world zones has a chance to drop this item.
This is used to donate to your guild and are used to level up your guild or to build buildings in your Guild Castle.
To donate Guild Runes:
- Press 'G' to display the Guild window
- Press the 'Resources' button, this will display the Resources window
- Open up your backpack, then drag and drop the Guild Rune into an empty slot in the Resources window
- Click on the 'Confirm' button
Note: I was playing this game for years before I realized that these are NOT Bound! You can mail them, share them, sell them in the AH... whatever!